Members of the crypto-zoological agency Monarch face off against a battery of god-sized monsters, including the mighty Godzilla, who collides with Mothra, Rodan, and his ultimate nemesis, the three-headed King Ghidorah. When these ancient super-species-thought to be mere myths-rise again, they all vie for supremacy, leaving humanity's very existence hanging in the balance.

When nuclear Armageddon destroys civilization on Earth, the only survivors are those on the 12 international space stations in orbit at the time. Three generations later, the 4,000 survivors living on a space ark of linked stations see their resources dwindle and face draconian measures established to ensure humanity's future. Desperately looking for a solution, the ark's leaders send 100 juvenile prisoners back to the planet to test its habitability. Having always lived in space, the exiles find the planet fascinating and terrifying, but with the fate of the human race in their hands, they must forge a path into the unknown.